Morphological and behavioral adaptations. Mechanisms of plant adaptation to unfavorable environmental conditions

a brown or green twig, and orthoptera insects imitate a leaf. Fish that lead a bottom-dwelling lifestyle (for example, flounder) have a flat body.

Protective coloration

Allows you to be invisible among the surrounding background. Thanks to the protective coloration, the organism becomes difficult to distinguish and, therefore, protected from predators. Bird eggs laid on sand or ground are gray and brown with spots, similar to the color of the surrounding soil. In cases where eggs are inaccessible to predators, they are usually colorless. Butterfly caterpillars are often green, the color of the leaves, or dark, the color of the bark or earth. Bottom fish are usually colored to match the color of the sandy bottom (rays and flounder). Moreover, flounders also have the ability to change color depending on the color of the surrounding background. The ability to change color by redistributing pigment in the integument of the body is also known in terrestrial animals (chameleon). Desert animals, as a rule, have a yellow-brown or sandy-yellow color. A monochromatic protective color is characteristic of both insects (locusts) and small lizards, as well as large ungulates (antelope) and predators (lion).

Warning coloring

Warns a potential enemy about the presence of defense mechanisms (presence toxic substances or special protection bodies). Warning coloring distinguishes poisonous, stinging animals and insects (snakes, wasps, bumblebees) from the environment with bright spots or stripes.

Mimicry

Mimicry is the result of homologous (identical) mutations in different species that help unprotected animals survive. For imitating species, it is important that their numbers are small compared to the model they are imitating, otherwise the enemies will not develop a stable negative reflex to the warning coloration. The low abundance of mimicking species is supported by a high concentration of lethal genes in the gene pool. When homozygous, these genes cause lethal mutations, resulting in a high percentage of individuals not surviving to adulthood.

Morphological adaptations involve changes in the shape or structure of an organism. An example of such an adaptation is a hard shell, which provides protection from predatory animals. Physiological adaptations are associated with chemical processes in the body. Thus, the smell of a flower can serve to attract insects and thereby contribute to pollination of the plant. Behavioral adaptation is associated with a certain aspect of an animal’s life. A typical example is winter dream at the bear. Most adaptations are a combination of these types. For example, blood sucking in mosquitoes is ensured by a complex combination of such adaptations as the development of specialized parts of the oral apparatus adapted to sucking, the formation of search behavior to find a prey animal, and the development salivary glands special secretions that prevent the clotting of sucked blood.

All plants and animals constantly adapt to their environment. To understand how this happens, it is necessary to consider not only the animal or plant as a whole, but also genetic basis adaptation.

Genetic basis.

In each species, the program for the development of traits is embedded in the genetic material. The material and the program encoded in it are passed on from one generation to the next, remaining relatively unchanged, so that representatives of a given species look and behave almost the same. However, in a population of organisms of any species there are always minor changes genetic material and, therefore, variation in the characteristics of individuals. It is from these diverse genetic variations that the process of adaptation selects those traits or favors the development of those traits that most increase the chances of survival and thereby the preservation of genetic material. Adaptation can thus be thought of as the process by which genetic material increases its chances of persistence in subsequent generations. From this point of view, each species represents successful way preservation of certain genetic material.

To pass on genetic material, an individual of any species must be able to feed, survive until the breeding season, leave offspring, and then spread them over as wide an area as possible.

Nutrition.

All plants and animals must receive energy and various substances from the environment, primarily oxygen, water and inorganic compounds. Almost all plants use the energy of the Sun, transforming it through the process of photosynthesis. Animals get energy by eating plants or other animals.

Each species is adapted in a certain way to provide itself with food. Hawks have sharp talons for capturing prey, and the location of the eyes in the front of the head allows them to judge the depth of space, which is necessary for hunting while flying at high speed. Other birds, such as herons, have developed long neck and legs. They obtain food by carefully wandering through shallow water and lying in wait for unwary aquatic animals. Darwin's finches, a group of closely related bird species from the Galapagos Islands, provide a classic example of highly specialized adaptation to in different ways nutrition. Thanks to certain adaptive morphological changes, primarily in the structure of the beak, some species became granivorous, others became insectivorous.

Turning to fish, predators such as sharks and barracudas have sharp teeth to catch prey. Others, such as small anchovies and herring, obtain small food particles by filtering seawater through comb-like gill rakers.

In mammals, an excellent example of adaptation to the type of nutrition is the structural features of teeth. The canines and molars of leopards and other felines are exceptionally sharp, which allows these animals to hold and tear the body of their prey. Deer, horses, antelopes and other grazing animals have large molars with wide, ribbed surfaces adapted for chewing grass and other plant foods.

Various ways to receive nutrients can be observed not only in animals, but also in plants. Many of them, primarily legumes - peas, clover and others - have developed symbiotic, i.e. mutually beneficial relationship with bacteria: bacteria convert atmospheric nitrogen into a chemical form available to plants, and plants provide energy to bacteria. Carnivorous plants such as sarracenia and sundew obtain nitrogen from the bodies of insects captured by trapping leaves.

Protection.

The environment consists of living and nonliving components. The living environment of any species includes animals that feed on members of that species. Adaptations of predatory species are aimed at efficient food acquisition; Prey species adapt to avoid becoming prey to predators.

Many potential prey species have protective or camouflage colors that hide them from predators. Thus, in some species of deer, the spotted skin of young individuals is invisible against the background of alternating spots of light and shadow, and white hares are difficult to distinguish against the background of snow cover. The long, thin bodies of stick insects are also difficult to see because they resemble twigs or twigs from bushes and trees.

Deer, hares, kangaroos and many other animals have evolved long legs that allow them to escape from predators. Some animals, such as opossums and hog snakes, have even developed a unique behavior called death faking, which increases their chances of survival, since many predators do not eat carrion.

Some types of plants are covered with thorns or thorns that repel animals. Many plants have a disgusting taste to animals.

Environmental factors, in particular climate, often place living organisms in difficult conditions. For example, animals and plants often have to adapt to temperature extremes. Animals escape the cold by using insulating fur or feathers, migrating to warmer climates, or hibernating. Most plants survive the cold by entering a state of dormancy, equivalent to hibernation in animals.

In hot weather, the animal cools itself by sweating or frequent breathing, which increases evaporation. Some animals, especially reptiles and amphibians, are able to enter summer hibernation, which is essentially similar to winter hibernation, but is caused by heat rather than cold. Others are simply looking for a cool place.

Plants can maintain their temperature to some extent by regulating the rate of evaporation, which has the same cooling effect as sweating in animals.

Reproduction.

A critical step in ensuring the continuity of life is reproduction, the process by which genetic material is passed on to the next generation. Reproduction has two important aspects: meeting of opposite-sex individuals to exchange genetic material and raise offspring.

Among the adaptations that ensure the meeting of individuals of different sexes is sound communication. In some species, the sense of smell plays an important role in this sense. For example, cats are strongly attracted to the smell of a cat in heat. Many insects secrete the so-called. attractants – chemical substances, attracting individuals of the opposite sex. Flower scents are an effective plant adaptation to attract pollinating insects. Some flowers smell sweet and attract nectar-feeding bees; others smell disgusting, attracting flies that feed on carrion.

Vision is also very important for meeting individuals of different sexes. In birds, the male's mating behavior, his lush feathers and bright colors attract the female and prepare her for copulation. Flower color in plants often indicates which animal is needed to pollinate that plant. For example, flowers pollinated by hummingbirds are colored red, which attracts these birds.

Many animals have developed ways to protect their offspring in the early stages of life. Most adaptations of this kind are behavioral and involve actions by one or both parents that increase the chances of survival of the young. Most birds build nests that are specific to each species. However, some species, such as the cowbird, lay eggs in the nests of other bird species and entrust the young to the parental care of the host species. In many birds and mammals, as well as some fish, there is a period when one of the parents takes great risks, taking on the function of protecting the offspring. Although this behavior sometimes threatens the death of the parent, it ensures the safety of the offspring and the preservation of genetic material.

A number of animal and plant species use a different reproductive strategy: they produce a huge number of offspring and leave them unprotected. In this case, the low chances of survival of an individual growing individual are balanced by the large number of offspring.

Settlement.

Most species have developed mechanisms to remove offspring from the places where they were born. This process, called dispersal, increases the likelihood that offspring will grow up in unoccupied territory.

Most animals simply avoid places where there is too much competition. However, evidence is accumulating that dispersal is driven by genetic mechanisms.

Many plants have adapted to dispersing seeds with the help of animals. Thus, the fruits of the cocklebur have hooks on the surface, with which they cling to the fur of passing animals. Other plants produce tasty, fleshy fruits, such as berries, that are eaten by animals; the seeds pass through the digestive tract and are “sown” intact elsewhere. Plants also use wind to spread. For example, the wind carries the “propellers” of maple seeds, as well as cottonweed seeds, which have tufts of fine hairs. Steppe plants such as tumbleweeds, which acquire a spherical shape by the time the seeds ripen, are driven by the wind over long distances, dispersing seeds along the way.

Above were just some of the most striking examples of adaptations. However, almost every trait of any species is the result of adaptation. All these signs form a harmonious combination, which allows the body to successfully lead its own special way of life. Man in all his features, from brain structure to shape thumb on the leg, is the result of adaptation. Adaptive traits contributed to the survival and reproduction of his ancestors, who had the same traits. In general, the concept of adaptation has great importance for all areas of biology.

Reactions to unfavorable environmental factors are detrimental to living organisms only under certain conditions, but in most cases they have adaptive significance. Therefore, these responses were called “general adaptation syndrome” by Selye. In later works, he used the terms “stress” and “general adaptation syndrome” as synonyms.

Adaptation is a genetically determined process of the formation of protective systems that ensure increased stability and the course of ontogenesis in unfavorable conditions for it.

Adaptation is one of the most important mechanisms that increases resilience biological system, including plant organisms, in changed conditions of existence. The better an organism is adapted to a certain factor, the more resistant it is to its fluctuations.

The genotypically determined ability of an organism to change metabolism within certain limits depending on the action of the external environment is called reaction norm. It is controlled by the genotype and is characteristic of all living organisms. Most modifications that occur within the normal range of reaction have adaptive significance. They correspond to changes in the environment and ensure better plant survival under fluctuating environmental conditions. In this regard, such modifications have evolutionary significance. The term “reaction norm” was introduced by V.L. Johannsen (1909).

The greater the ability of a species or variety to be modified in accordance with the environment, the wider its reaction rate and the higher its ability to adapt. This property distinguishes resistant varieties of crops. As a rule, slight and short-term changes in environmental factors do not lead to significant disturbances in the physiological functions of plants. This is due to their ability to maintain relative dynamic balance of the internal environment and the stability of basic physiological functions in a changing external environment. At the same time, sudden and prolonged impacts lead to disruption of many functions of the plant, and often to its death.

Adaptation includes all processes and adaptations (anatomical, morphological, physiological, behavioral, etc.) that contribute to increased stability and contribute to the survival of the species.

1.Anatomical and morphological devices. In some representatives of xerophytes, the length of the root system reaches several tens of meters, which allows the plant to use groundwater and not experience a lack of moisture in conditions of soil and atmospheric drought. In other xerophytes, the presence of a thick cuticle, pubescent leaves, and the transformation of leaves into spines reduce water loss, which is very important in conditions of lack of moisture.

Stinging hairs and spines protect plants from being eaten by animals.

Trees in the tundra or at high mountain altitudes look like squat creeping shrubs; in winter they are covered with snow, which protects them from severe frosts.

In mountainous regions with large daily temperature fluctuations, plants often have the form of spread out pillows with numerous stems densely spaced. This allows you to maintain moisture inside the pillows and a relatively uniform temperature throughout the day.

In the swamp and aquatic plants a special air-bearing parenchyma (aerenchyma) is formed, which is an air reservoir and facilitates the breathing of plant parts immersed in water.

2. Physiological-biochemical adaptations. In succulents, an adaptation for growing in desert and semi-desert conditions is the assimilation of CO 2 during photosynthesis via the CAM pathway. These plants have stomata that are closed during the day. Thus, the plant preserves its internal water reserves from evaporation. In deserts, water is the main factor limiting plant growth. The stomata open at night, and at this time CO 2 enters the photosynthetic tissues. The subsequent involvement of CO 2 in the photosynthetic cycle occurs during the day when the stomata are closed.

Physiological and biochemical adaptations include the ability of stomata to open and close, depending on external conditions. Synthesis in cells of abscisic acid, proline, protective proteins, phytoalexins, phytoncides, increased activity of enzymes that counteract oxidative breakdown organic matter, accumulation of sugars in cells and a number of other changes in metabolism help to increase plant resistance to unfavorable conditions external environment.

The same biochemical reaction can be carried out by several molecular forms of the same enzyme (isoenzymes), with each isoform exhibiting catalytic activity in a relatively narrow range of some environmental parameter, such as temperature. The presence of a number of isoenzymes allows the plant to carry out reactions in a much wider temperature range compared to each individual isoenzyme. This allows the plant to successfully perform vital functions in changing temperature conditions.

3. Behavioral adaptations, or avoidance of an unfavorable factor. An example is ephemera and ephemeroids (poppy, chickweed, crocuses, tulips, snowdrops). They go through their entire development cycle in the spring in 1.5-2 months, even before the onset of heat and drought. Thus, they seem to leave, or avoid falling under the influence of the stressor. Similarly, early ripening varieties of agricultural crops form a harvest before the onset of unfavorable seasonal phenomena: August fogs, rains, frosts. Therefore, the selection of many agricultural crops is aimed at creating early ripening varieties. Perennial plants overwinter in the form of rhizomes and bulbs in the soil under snow, which protects them from freezing.

Adaptation of plants to unfavorable factors is carried out simultaneously at many levels of regulation - from an individual cell to a phytocenosis. The higher the level of organization (cell, organism, population), the larger number mechanisms simultaneously participates in the adaptation of plants to stress.

Regulation of metabolic and adaptation processes inside the cell is carried out using systems: metabolic (enzymatic); genetic; membrane These systems are closely interconnected. Thus, the properties of membranes depend on gene activity, and the differential activity of the genes themselves is under the control of membranes. The synthesis of enzymes and their activity are controlled at the genetic level, while at the same time enzymes regulate nucleic acid metabolism in the cell.

On organismal level new ones are added to the cellular mechanisms of adaptation, reflecting the interaction of organs. In unfavorable conditions, plants create and retain such an amount of fruit elements that are sufficiently provided with the necessary substances to form full-fledged seeds. For example, in the inflorescences of cultivated cereals and in the crowns fruit trees in unfavorable conditions, more than half of the established ovaries may fall off. Such changes are based on competitive relations between organs for physiologically active and nutrients.

Under stress conditions, the processes of aging and falling of the lower leaves sharply accelerate. At the same time, substances needed by plants move from them to young organs, responding to the organism’s survival strategy. Thanks to the recycling of nutrients from the lower leaves, the younger ones, the upper leaves, remain viable.

Mechanisms for regeneration of lost organs operate. For example, the surface of a wound is covered with secondary integumentary tissue (wound periderm), a wound on a trunk or branch is healed with nodules (calluses). When the apical shoot is lost, dormant buds awaken in plants and side shoots intensively develop. The regeneration of leaves in the spring instead of those that fell in the fall is also an example of natural organ regeneration. Regeneration as a biological adaptation that provides vegetative propagation plants by segments of root, rhizome, thallus, stem and leaf cuttings, isolated cells, individual protoplasts, has a large practical significance for plant growing, fruit growing, forestry, ornamental gardening, etc.

The hormonal system also participates in the processes of protection and adaptation at the plant level. For example, under the influence of unfavorable conditions in a plant, the content of growth inhibitors sharply increases: ethylene and abscisic acid. They reduce metabolism, inhibit growth processes, accelerate aging, organ loss, and the plant’s transition to a dormant state. Inhibition of functional activity under stress conditions under the influence of growth inhibitors is a characteristic reaction for plants. At the same time, the content of growth stimulants in tissues decreases: cytokinin, auxin and gibberellins.

On population level selection is added, which leads to the emergence of more adapted organisms. The possibility of selection is determined by the existence of intrapopulation variability in plant resistance to various factors external environment. An example of intrapopulation variability in resistance can be the uneven emergence of seedlings on saline soil and the increase in variation in germination timing with increasing stressors.

View in modern concept consists of a large number of biotypes - smaller ecological units that are genetically identical, but exhibit different resistance to environmental factors. Under different conditions, not all biotypes are equally viable, and as a result of competition, only those that best meet the given conditions remain. That is, the resistance of a population (variety) to one or another factor is determined by the resistance of the organisms that make up the population. Resistant varieties include a set of biotypes that provide good productivity even in unfavorable conditions.

At the same time, during long-term cultivation of varieties, the composition and ratio of biotypes in the population changes, which affects the productivity and quality of the variety, often not for the better.

So, adaptation includes all processes and adaptations that increase the resistance of plants to unfavorable environmental conditions (anatomical, morphological, physiological, biochemical, behavioral, population, etc.)

But to choose the most effective adaptation path, the main thing is the time during which the body must adapt to new conditions.

In the event of a sudden action of an extreme factor, the response cannot be delayed; it must follow immediately to avoid irreversible damage to the plant. With prolonged exposure to a small force, adaptive changes occur gradually, and the choice of possible strategies increases.

In this regard, there are three main adaptation strategies: evolutionary, ontogenetic And urgent. The goal of the strategy is the effective use of available resources to achieve the main goal - the survival of the body under stress. The adaptation strategy is aimed at maintaining the structural integrity of vital macromolecules and the functional activity of cellular structures, preserving life regulation systems, and providing plants with energy.

Evolutionary or phylogenetic adaptations(phylogeny - development biological species in time) are adaptations that arise during the evolutionary process based on genetic mutations, selection and are inherited. They are the most reliable for plant survival.

In the process of evolution, each plant species has developed certain needs for living conditions and adaptability to the ecological niche it occupies, a stable adaptation of the organism to its habitat. Moisture and shade tolerance, heat resistance, cold resistance and other ecological characteristics of specific plant species were formed as a result of long-term exposure to appropriate conditions. Thus, heat-loving and short-day plants are characteristic of southern latitudes, less demanding of heat and long-day plants - for northern ones. Numerous evolutionary adaptations of xerophyte plants to drought are well known: economical use of water, deep-lying root system, shedding leaves and transition to a dormant state, and other adaptations.

In this regard, varieties of agricultural plants exhibit resistance precisely to those environmental factors against the background of which breeding and selection of productive forms is carried out. If selection takes place in a number of successive generations against the background of the constant influence of some unfavorable factor, then the resistance of the variety to it can be significantly increased. It is natural that the varieties selected by the research institute Agriculture South-East (Saratov), ​​are more resistant to drought than varieties created in breeding centers of the Moscow region. In the same way in ecological zones With unfavorable soil-climatic conditions, resistant local plant varieties have formed, and endemic plant species are resistant precisely to the stressor that is expressed in their habitat.

Characteristics of resistance of spring wheat varieties from the collection of the All-Russian Institute of Plant Growing (Semyonov et al., 2005)

In a natural setting, environmental conditions usually change very quickly, and the time during which the stress factor reaches a damaging level is not enough for the formation of evolutionary adaptations. In these cases, plants use not permanent, but stressor-induced defense mechanisms, the formation of which is genetically predetermined (determined).

Ontogenetic (phenotypic) adaptations not related to genetic mutations and are not inherited. The formation of this kind of adaptation takes a relatively long time, which is why they are called long-term adaptations. One of these mechanisms is the ability of a number of plants to form a water-saving CAM-type photosynthetic pathway under conditions of water deficiency caused by drought, salinity, low temperatures and other stressors.

This adaptation is associated with the induction of the expression of the phosphoenolpyruvate carboxylase gene, which is “inactive” under normal conditions, and the genes of other enzymes of the CAM pathway of CO 2 assimilation, with the biosynthesis of osmolytes (proline), with the activation of antioxidant systems and changes in the daily rhythms of stomatal movements. All this leads to very economical use of water.

In field crops, for example, corn, aerenchyma is absent under normal growing conditions. But under conditions of flooding and a lack of oxygen in the tissues of the roots, some of the cells of the primary cortex of the root and stem die (apoptosis, or programmed cell death). In their place, cavities are formed through which oxygen is transported from the aboveground part of the plant to the root system. The signal for cell death is ethylene synthesis.

Urgent adaptation occurs with rapid and intense changes in living conditions. It is based on the formation and functioning of shock defense systems. Shock defense systems include, for example, the heat shock protein system, which is formed in response to a rapid increase in temperature. These mechanisms provide short-term conditions for survival under the influence of a damaging factor and thereby create the prerequisites for the formation of more reliable long-term specialized adaptation mechanisms. An example of specialized adaptation mechanisms is the new formation of antifreeze proteins at low temperatures or the synthesis of sugars during the overwintering of winter crops. At the same time, if the damaging effect of a factor exceeds the protective and reparation capabilities of the body, then death inevitably occurs. In this case, the organism dies at the stage of urgent or at the stage of specialized adaptation, depending on the intensity and duration of the extreme factor.

Distinguish specific And nonspecific (general) plant responses to stressors.

Nonspecific reactions do not depend on the nature of the acting factor. They are the same under the influence of high and low temperatures, lack or excess of moisture, high concentration of salts in the soil or harmful gases in the air. In all cases, the permeability of membranes in plant cells increases, respiration is impaired, the hydrolytic breakdown of substances increases, the synthesis of ethylene and abscisic acid increases, and cell division and elongation are inhibited.

The table presents a complex of nonspecific changes that occur in plants under the influence of various environmental factors.

Changes in physiological parameters in plants under the influence of stress conditions (according to G.V. Udovenko, 1995)

Based on the data in the table, it can be seen that plant resistance to several factors is accompanied by unidirectional physiological changes. This gives reason to believe that an increase in plant resistance to one factor may be accompanied by an increase in resistance to another. This has been confirmed by experiments.

Experiments at the Institute of Plant Physiology of the Russian Academy of Sciences (Vl. V. Kuznetsov and others) have shown that short-term heat treatment of cotton plants is accompanied by an increase in their resistance to subsequent salinity. And the adaptation of plants to salinity leads to an increase in their resistance to high temperatures. Heat shock increases the ability of plants to adapt to subsequent drought and, conversely, during drought the body's resistance to high temperatures increases. Short-term exposure to high temperatures increases resistance to heavy metals and UV-B irradiation. Previous drought promotes plant survival in salinity or cold conditions.

The process of increasing the body's resistance to this environmental factor as a result of adaptation to a factor of a different nature is called cross adaptation.

To study general (nonspecific) mechanisms of resistance, the response of plants to factors that cause water deficit: for salinity, drought, low and high temperatures and some others. At the level of the whole organism, all plants respond to water deficiency in the same way. Characterized by inhibition of shoot growth, increased growth of the root system, abscisic acid synthesis, and decreased stomatal conductance. After some time, the lower leaves age rapidly and their death is observed. All these reactions are aimed at reducing water consumption by reducing the evaporating surface, as well as by increasing the absorption activity of the root.

Specific reactions- These are reactions to the action of any one stress factor. Thus, phytoalexins (substances with antibiotic properties) are synthesized in plants in response to contact with pathogens.

The specificity or non-specificity of response reactions implies, on the one hand, the attitude of the plant to various stressors and, on the other hand, the specificity of the reactions of plants of different species and varieties to the same stressor.

The manifestation of specific and nonspecific plant responses depends on the strength of stress and the speed of its development. Specific responses occur more often if stress develops slowly, and the body has time to rebuild and adapt to it. Nonspecific reactions usually occur with a more short-term and strong action stressor The functioning of nonspecific (general) resistance mechanisms allows the plant to avoid large energy expenditures for the formation of specialized (specific) adaptation mechanisms in response to any deviation from the norm in their living conditions.

Plant resistance to stress depends on the phase of ontogenesis. The most stable plants and plant organs are in a dormant state: in the form of seeds, bulbs; woody perennials - in a state of deep dormancy after leaf fall. Plants are most sensitive at a young age, since under stress conditions growth processes are damaged first. The second critical period is the period of gamete formation and fertilization. Stress during this period leads to a decrease in the reproductive function of plants and a decrease in yield.

If stressful conditions are repeated and have low intensity, then they contribute to plant hardening. This is the basis for methods of increasing resistance to low temperatures, heat, salinity, and increased levels of harmful gases in the air.

Reliability of a plant organism is determined by its ability to prevent or eliminate failures in different levels biological organization: molecular, subcellular, cellular, tissue, organ, organismal and population.

To prevent disruptions in plant life under the influence of unfavorable factors, the principles of redundancy, heterogeneity of functionally equivalent components, systems for repairing lost structures.

Redundancy of structures and functionality is one of the main ways to ensure system reliability. Redundancy and redundancy have diverse manifestations. At the subcellular level, the redundancy and duplication of genetic material contribute to increasing the reliability of the plant organism. This is ensured, for example, by the double helix of DNA and an increase in ploidy. The reliability of the functioning of a plant organism under changing conditions is also supported by the presence of various messenger RNA molecules and the formation of heterogeneous polypeptides. These include isoenzymes that catalyze the same reaction, but differ in their physicochemical properties and the stability of the molecular structure under changing environmental conditions.

At the cellular level, an example of redundancy is an excess of cellular organelles. Thus, it has been established that a portion of the available chloroplasts is sufficient to provide the plant with photosynthetic products. The remaining chloroplasts seem to remain in reserve. The same applies to the total chlorophyll content. Redundancy is also manifested in the large accumulation of precursors for the biosynthesis of many compounds.

At the organismal level, the principle of redundancy is expressed in the formation and in the laying down at different times of more than is required for the change of generations, the number of shoots, flowers, spikelets, in a huge amount of pollen, ovules, and seeds.

At the population level, the principle of redundancy is manifested in a large number of individuals that differ in resistance to a particular stress factor.

Reparation systems also operate at different levels - molecular, cellular, organismal, population and biocenotic. Repair processes require energy and plastic substances, so repair is possible only if sufficient metabolic rate is maintained. If metabolism stops, repair also stops. In extreme environmental conditions, maintaining respiration is especially important, since it is respiration that provides energy for repair processes.

The restorative ability of cells of adapted organisms is determined by the resistance of their proteins to denaturation, namely the stability of the bonds that determine the secondary, tertiary and quaternary structure of the protein. For example, the resistance of mature seeds to high temperatures, as a rule, is due to the fact that after dehydration their proteins become resistant to denaturation.

The main source of energy material as a substrate for respiration is photosynthesis, therefore, the energy supply of the cell and the associated repair processes depend on the stability and ability of the photosynthetic apparatus to recover after damage. To maintain photosynthesis under extreme conditions in plants, the synthesis of thylakoid membrane components is activated, lipid oxidation is inhibited, and the ultrastructure of plastids is restored.

At the organismal level, an example of regeneration can be the development of replacement shoots, the awakening of dormant buds when growth points are damaged.

If you find an error, please highlight a piece of text and click Ctrl+Enter.

To survive in unfavorable climatic conditions, plants, animals and birds have some features. These features are called "physiological adaptations", examples of which can be seen in almost every species of mammal, including humans.

Why is physiological adaptation necessary?

Living conditions in some parts of the planet are not entirely comfortable, nevertheless, there are various representatives of wildlife there. There are several reasons why these animals did not leave the unfavorable environment.

First of all, climatic conditions may have changed when a certain species already existed in a given area. Some animals are not adapted to migration. It is also possible that territorial features do not allow migration (islands, mountain plateaus, etc.). For a certain species, changed habitat conditions still remain more suitable than in any other place. AND physiological adaptation is the best option solving the problem.

What do you mean by adaptation?

Physiological adaptation is the harmony of organisms with a specific habitat. For example, the comfortable stay of its inhabitants in the desert is due to their adaptation to high temperatures and lack of access to water. Adaptation is the appearance of certain characteristics in organisms that allow them to get along with some elements of the environment. They arise during the process of certain mutations in the body. Physiological adaptations, examples of which are well known in the world, are, for example, the ability to echolocation in some animals (bats, dolphins, owls). This ability helps them navigate in a space with limited lighting (in the dark, in water).

Physiological adaptation is a set of reactions of the body to certain pathogenic factors in the environment. It provides organisms with a greater likelihood of survival and is one of the methods of natural selection for strong and resilient organisms in a population.

Types of physiological adaptation

Adaptation of the organism is distinguished between genotypic and phenotypic. Genotypic is based on the conditions of natural selection and mutations that led to changes in organisms of an entire species or population. It was in the process of this type of adaptation that modern species of animals, birds and humans were formed. The genotypic form of adaptation is hereditary.

The phenotypic form of adaptation is due to individual changes in a particular organism for a comfortable stay in certain climatic conditions. It can also develop due to constant exposure to an aggressive environment. As a result, the body acquires resistance to its conditions.

Complex and cross adaptations

Complex adaptations occur in certain climatic conditions. For example, the body gets used to low temperatures during a long stay in the northern regions. This form of adaptation develops in every person when moving to a different climate zone. Depending on the characteristics of a particular organism and its health, this form of adaptation proceeds in different ways.

Cross adaptation is a form of habituation of the organism in which the development of resistance to one factor increases resistance to all factors of this group. A person's physiological adaptation to stress increases his resistance to some other factors, for example, to cold.

Based on positive cross adaptations A set of measures has been developed to strengthen the heart muscle and prevent heart attacks. IN natural conditions those people who most often in life have encountered stressful situations, are less susceptible to the consequences of myocardial infarction than those who led a quiet lifestyle.

Types of adaptive reactions

There are two types of adaptive reactions of the body. The first type is called “passive adaptations”. These reactions take place at the cellular level. They characterize the formation of the degree of resistance of the body to the effects of negative environmental factors. For example, change atmospheric pressure. Passive adaptation allows you to maintain the normal functionality of the body with small fluctuations in atmospheric pressure.

The most well-known physiological adaptations in passive animals are defensive reactions living organism exposed to cold. Hibernation, in which life processes slow down, is inherent in some species of plants and animals.

The second type of adaptive reactions is called active and involves the body’s protective measures when exposed to pathogenic factors. In this case, the internal environment of the body remains constant. This type of adaptation is characteristic of highly developed mammals and humans.

Examples of physiological adaptations

Physiological adaptation of a person is manifested in all situations that are non-standard for his environment and lifestyle. Acclimatization is the most famous example of adaptation. For different organisms this process occurs at different speeds. Some people need a few days to get used to new conditions, for many it will take months. Also, the speed of adaptation depends on the degree of difference from the usual habitat.

In hostile environments, many mammals and birds have a characteristic set of body responses that make up their physiological adaptations. Examples (in animals) can be observed in almost every climate zone. For example, desert dwellers accumulate reserves of subcutaneous fat, which oxidizes and forms water. This process is observed before the onset of a period of drought.

Physiological adaptation in plants also takes place. But it is passive in nature. An example of such an adaptation is trees shedding leaves when the cold season approaches. The bud areas are covered with scales, which protect them from the harmful effects of low temperatures and snow and wind. Metabolic processes in plants slow down.

In combination with morphological adaptation, the physiological reactions of the body provide it with a high level of survival in unfavorable conditions and during sudden changes in the environment.